Witnesses report air strikes and paramilitary forces fire anti-aircraft weapons while WHO warns of "huge biological risk" after fighters occupy laboratory in capital Khartoum.
Sudan's army chief Abdel Fattah al Burhan says there was "no room" for negotiations with his rival paramilitary commander Mohamed Hamdan Dagalo as fighting rages and death toll tops 330.
Fighting continues despite a 24-hour ceasefire lobbied for by countries trying to evacuate their citizens as conflict leaves nearly 300 people dead and thousands wounded.
The eruption of violence follows tensions between military leader Abdel Fattah al Burhan and paramilitary commander Mohamed Hamdan Daglo over the planned integration of paramilitary forces into the regular army.
Army says the deployment of Rapid Support Forces in the capital “stirred up panic and fear among people" and exacerbated security risks while increasing tensions with regular forces.
Huge crowds blocked main roads and marched in Sudan's several cities, facing heavy tear gas fired by the security forces and many were seen breaking their Ramadan fasts in the street.
Protester died of wounds sustained after he was "hit in the abdomen by a bullet fired by security forces," says Central Committee of Sudan Doctors, taking total death toll from military crackdown on demonstrations to 120.
At least 118 people have been killed while demanding a return to civilian rule, which is a condition set by Western governments to resume crucial aid, currently halted as a result of the military coup.
It was not immediately clear what started the fighting between members of rival ethnic Nuba and Misseriya people in the town of Lagawa.
Witnesses say they heard shooting and that houses had been burnt in Blue Nile state, where seven people were killed and over 20 suffered injuries.
Karim Khan is in Sudan a year after he visited the country for talks on outstanding arrest warrants over crimes committed during the 2003 Darfur war under ousted president Omar al Bashir.
Witnesses say security forces were trying to prevent swelling numbers of demonstrators from marching towards the presidential palace, the military's seat of power in the heart of the capital city.
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